for they fail to recognize that in addition to gene1.fail to 2.in addition to

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for they fail to recognize that in addition to gene1.fail to 2.in addition to
for they fail to recognize that in addition to gene
1.fail to
2.in addition to

for they fail to recognize that in addition to gene1.fail to 2.in addition to
1.没能...
2.另外考虑到...

没有
除了